Surly the squirrel and his animal friends enjoy a peaceful life in their urban park after the events that originally united them. Unfortunately, their harmonious existence is threatened when the corrupt mayor of Oakton City announces plans to destroy the park and replace it with an amusement park. Surly, with his trusted friend Buddy the rat, rallies his fellow creatures, including the precocious mouse Andie and the loyal pug Precious, to thwart the mayor's scheme and preserve their home.

Faced with an uphill battle against the mayor's ambitious construction project, the group devises clever tactics to disrupt the progress. Meanwhile, they encounter unexpected allies and face formidable adversaries, all while learning the true meaning of friendship and community. As the conflict intensifies, Surly and his friends must rely on their unity, resourcefulness, and determination to save their beloved park and ensure a safer future for all its inhabitants.